Published papers

A list of scientific publications on Ethiopian wolves, many authored/co-authored by EWCP staff


Atickem, A and Stenseth N.Chr. (2022) The role of rodents in the conservation of endangered species in the Ethiopian highlands. Therya, 13:73-77 doi:10.12933/therya-22-1185


Krofel, M., Hatlauf, J., Bogdanowicz, W., Campbell, L. A. D., Godinho, R., Jhala, Y. V., Kitchener, A. C., Koepfli, K.-P., Moehlman, P., Senn, H., Sillero-Zubiri, C., Viranta, S., Werhahn, G., & Alvares, F. (2021). Towards resolving taxonomic uncertainties in wolf, dog and jackal lineages of Africa, Eurasia and Australasia. Journal of Zoology, 00, 1– 14.


Merkebu S and Yazezew D. 2021. Assessment of human-wildlife conflict and the attitude of local communities to wild animal conservation around Borena Sayint National Park, Ethiopia. International Journal of Ecology, 2021 Article ID 6619757.


Šklíba J, Vlasatá T, Lövy M, Hrouzková E, Meheretu Y, Sillero‐Zubiri C and Šumbera R. 2020. The giant that makes do with little: small and easy‐to‐leave home ranges found in the giant root‐rat. Journal of Zoology, 310: 64-70. doi:10.1111/jzo.12729


Hrouzková E, Bernasová E and Šklíba J. 2020. Eavesdropping on a heterospecific alarm call in the giant root-rat (Tachyorytes macrocephalus), an important prey of the Ethiopian wolf (Canis simensis). Journal of Ethology 38: 121–124.


Estifanos, T.K., et al., Managing conflicts between local land use and the protection of the Ethiopian wolf: Residents’ preferences for conservation program design features. Ecological Economics, 2020. 169: p. 15. DOI: 10.1016/j.ecolecon.2019.106511

Foley, C.J., C. Sillero‐Zubiri, and L. Börger, Opensource, lowcost modular GPS collars for monitoring and tracking wildlife. Methods in Ecology and Evolution, 2020. 11(4): p. 553-558.

Estifanos, T., et al., What are tourists willing to pay for securing the survival of a flagship species? The case of protection of the Ethiopian wolf. Tourism Economics, 2019.

Gutema, T.M., et al., Foraging ecology of African wolves (Canis lupaster) and its implications for the conservation of Ethiopian wolves (Canis simensis). R Soc Open Sci, 2019. 6(9): p. 190772.

Gopalakrishnan, S., et al., Interspecific Gene Flow Shaped the Evolution of the Genus Canis. Curr Biol, 2019. 29(23): p. 4152.

Macdonald, D.W., et al., Monogamy: Cause, Consequence, or Corollary of Success in Wild Canids? Frontiers in Ecology and Evolution, 2019. 7: p. 28.

Eshete, G., et al., Does livestock predation reflect in negative local perceptions of Ethiopian wolves in South Wollo? Tropical Ecology, 2018. 59(1): p. 11-19. Link

Eshete, G., J. Marino, and C. Sillero-Zubiri, Ethiopian wolves conflict with pastoralists in small Afroalpine relicts. African Journal of Ecology, 2018. 56(2): p. 368-374.


Gutema, T.M., et al., Competition between sympatric wolf taxa: an example involving African and Ethiopian wolves. R Soc Open Sci, 2018. 5(5): p. 172207.


Perry, L.R., J. Marino, and C. Sillero-Zubiri, Going to the Dogs: Free-Ranging Domestic Dogs Threaten an Endangered Wild Canid through Competitive Interactions. Journal of Biodiversity & Endangered Species, 2018. 6(1).


Gebresenbet, F., et al., A culture of tolerance: coexisting with large carnivores in the Kafa Highlands, Ethiopia. Oryx, 2017. 52(4): p. 751-760.


Marston, D.A., et al., Complete Genomic Sequence of Canine Distemper Virus from an Ethiopian Wolf. Genome Announc, 2017. 5(29): p. 2.

Macdonald, D.W., Animal behaviour and its role in carnivore conservation: examples of seven deadly threats. Animal Behaviour, 2016. 120: p. 197-209.


Marino, J., et al., Rabies and Distemper Outbreaks in Smallest Ethiopian Wolf Population. Emerg Infect Dis, 2017. 23(12): p. 2102-2104.


Vlasatá, T., et al., Daily activity patterns in the giant root rat (Tachyoryctes macrocephalus), a fossorial rodent from the Afro-alpine zone of the Bale Mountains, Ethiopia. Journal of Zoology, 2017. 302(3): p. 157-163.


Sillero-Zubiri, C., et al., Feasibility and efficacy of oral rabies vaccine SAG2 in endangered Ethiopian wolves. Vaccine, 2016. 34(40): p. 4792-8.


Eshete, G., et al., Community Resource Uses and Ethiopian Wolf Conservation in Mount Abune Yosef. Environ Manage, 2015. 56(3): p. 684-94.


Gordon, C.H., et al., Canine distemper in endangered Ethiopian wolves. Emerg Infect Dis, 2015. 21(5): p. 824-32.


Marston, D.A., et al., Complete genomic sequence of rabies virus from an ethiopian wolf. Genome Announc, 2015. 3(2): p. 2.


van Kesteren, F., et al., Helminth parasites in the endangered Ethiopian wolf, Canis simensis. J Helminthol, 2015. 89(4): p. 487-95.


Venkataraman, V.V., et al., Solitary Ethiopian wolves increase predation success on rodents when among grazing gelada monkey herds. Journal of Mammalogy, 2015. 96(1): p. 129-137.


Venkataraman, V.V., et al., Gelada herds increase predation success of Ethiopian wolves on rodents. American Journal of Physical Anthropology, 2014. 153: p. 260-260.


Yihune, M. and A. Bekele, Feeding ecology of the Ethiopian wolf in the Simien Mountains National Park, Ethiopia. African Journal of Ecology, 2014. 52(4): p. 484-490.


Gottelli, D., et al., Genetic structure and patterns of gene flow among populations of the endangered Ethiopian wolf. Animal Conservation, 2013. 16(2): p. 234-247. 


Marino, J., et al., The fall and rise of Ethiopian wolves: lessons for conservation of long-lived, social predators. Animal Conservation 2013: p. DOI: 10.1111/acv.12036.


van Kesteren, F., et al., The physiology of cooperative breeding in a rare social canid; sex, suppression and pseudopregnancy in female Ethiopian wolves. Physiol Behav, 2013. 122: p. 39-45. 


Marino, J., et al., Ecological bases of philopatry and cooperation in Ethiopian wolves. Behavioral Ecology and Sociobiology, 2012. 66(7): p. 1005-1015. Link 

Tallents, L.A., et al., Territory quality determines social group composition in Ethiopian wolves Canis simensis. J Anim Ecol, 2012. 81(1): p. 24-35.


van Kesteren, F., et al., Sex, stress and social status: patterns in fecal testosterone and glucocorticoid metabolites in male Ethiopian wolves. Gen Comp Endocrinol, 2012. 179(1): p. 30-7.


Kennedy, L.J., et al., Major histocompatibility complex diversity in the endangered Ethiopian wolf (Canis simensis). Tissue Antigens, 2011. 77(2): p. 118-25.


Sillero-Zubiri, C., et al., Ecology and reproductive strategy of an Afroalpine specialist: Ethiopian wolves in the Bale Mountains Walia Special Edition on the Bale Mountains, 2011: p. 61-79. Link


Fashing, P.J., N. Nguyen, and N.J. Fashing, Behavior of geladas and other endemic wildlife during a desert locust outbreak at Guassa, Ethiopia: ecological and conservation implications. Primates, 2010. 51(3): p. 193-7. DOI: 10.1007/s10329-010-0194-6


Hoffmann, M., et al., The impact of conservation on the status of the world's vertebrates. Science, 2010. 330(6010): p. 1503-9. DOI: 10.1126/science.1194442


Johnson, N., et al., A new outbreak of rabies in rare Ethiopian wolves (Canis simensis). Arch Virol, 2010. 155(7): p. 1175-7.


Tessema, M.E., et al., Community Attitudes Toward Wildlife and Protected Areas in Ethiopia. Society & Natural Resources, 2010. 23(6): p. 489-506.


Vial, F., et al., An analysis of long-term trends in the abundance of domestic livestock and free-roaming dogs in the Bale Mountains National Park. African Journal of Ecology, 2010.


Atickem, A., A. Bekele, and S.D. Williams, Competition between domestic dogs and Ethiopian wolf in the Bale Mountains National Park, Ethiopia. African Journal of Ecology, 2009. 48: p. 401-407.


Evangelista, P., R. Engeman, and L. Tallents, Testing a passive tracking index for monitoring the endangered Ethiopian wolf. Integr Zool, 2009. 4(2): p. 172-178.


Marino, J., R. Mitchell, and P.J. Johnson, Dietary specialization and climatic-linked variations in extant populations of Ethiopian wolves. African Journal of Ecology, 2009. 48(2): p. 517-525.


Thirgood, S., New perspectives on managing wildlife diseases. Journal of Applied Ecology, 2009. 46(2): p. 454-456.


Randall, D.A., et al., Fine-scale genetic structure in Ethiopian wolves imposed by sociality, migration, and population bottlenecks. Conservation Genetics, 2009. 11(1): p. 89-101. 

Schipper, J., et al., The status of the world's land and marine mammals: diversity, threat, and knowledge. Science, 2008. 322(5899): p. 225-30.


Yihune, M., A. Bekele, and Z. Tefera, Human-Ethiopian wolf conflict in and around the Simien Mountains National Park, Ethiopia. International Journal of Ecology and Environmental Sciences, 2008. 34(2): p. 149-155. DOI: 10.4314/sinet.v32i1.67785


Knobel, D.L., et al., Trapping and vaccination of endangered Ethiopian wolves to control an outbreak of rabies. Journal of Applied Ecology, 2007. 45(1): p. 109-116.


Randall, D.A., et al., Inbreeding is reduced by female-biased dispersal and mating behavior in Ethiopian wolves. Behavioral Ecology, 2007. 18(3): p. 579-589.


Haydon, D.T., et al., Low-coverage vaccination strategies for the conservation of endangered species. Nature, 2006. 443(7112): p. 692-5.


Marino, J., C. Sillero-Zubiri, and D.W. Macdonald, Trends, dynamics and resilience of an Ethiopian wolf population. Animal Conservation, 2006. 9(1): p. 49-58. DOI: 10.1111/j.1469-1795.2005.00011.x


Randall, D.A., et al., An integrated disease management strategy for the control of rabies in Ethiopian wolves. Biological Conservation, 2006. 131(2): p. 151-162.


Ashenafi, Z.T., Coulson, T., Sillero‐Zubiri, C. and Leader‐Williams, N. (2005), Behaviour and ecology of the Ethiopian wolf (Canis simensis) in a human‐dominated landscape outside protected areas. Animal Conservation, 8: 113-121.


Gottelli, D., et al., The effect of the last glacial age on speciation and population genetic structure of the endangered Ethiopian wolf (Canis simensis). Mol Ecol, 2004. 13(8): p. 2275-86.


Randall, D.A., et al., Rabies in endangered Ethiopian wolves. Emerg Infect Dis, 2004. 10(12): p. 2214-7.


Marino, J., Threatened Ethiopian wolves persist in small isolated Afroalpine enclaves. Oryx, 2003. 37(01): p. 62-71.


Edinburgh graduate helps protect Ethiopian wolves. Journal of Small Animal Practice, 2003. 44(1): p. 51-52.


Haydon, D.T., M.K. Laurenson, and C. Sillero-Zubiri, Integrating epidemiology into population viability analysis: Managing the risk posed by rabies and canine distemper to the Ethiopian wolf. Conservation Biology, 2002. 16(5): p. 1372 1385.


Stephens, P.A., et al., Impact of livestock and settlement on the large mammalian wildlife of Bale Mountains National Park, southern Ethiopia. Biological Conservation, 2001. 100: p. 307-322. DOI: 10.1016/S0006-3207(01)00035-0

Laurenson, M.K., et al., Disease threats to endangered species: patterns of infection by canine pathogens in Ethiopian wolves (Canis simensis) and sympatric domestic dogs. Animal Conservation, 1998. 1: p.273-280.


Sillero-Zubiri, C. and D.W. Macdonald, Scent-marking and territorial behaviour of Ethiopian wolves Canis simensis. Journal of Zoology, 1998. 245(3): p. 351-361.


The Ethiopian wolf: An action plan for its conservation. Oryx, 1998. 32(3): p. 174-175.


Whitby, J.E., P. Johnstone, and C. Sillero-Zubiri, Rabies virus in the decomposed brain of an Ethiopian wolf detected by nested reverse transcription-polymerase chain reaction. J Wildl Dis, 1997. 33(4): p. 912-5.


Sillero-Zubiri, C., P.J. Johnson, and D.W. MacDonald, A Hypothesis for Breeding Synchrony in Ethiopian Wolves (Canis simensis). Journal of Mammalogy, 1998. 79(3): p. 853-858.


Malcolm, J., The diet of the Ethiopian wolf (Canis simensis Ruppell) from a grassland are of the Bale Mountains, Ethiopia. African Journal of Ecology, 1997. 35(2): p. 162-164.


Sillero-Zubiri, C., Field immobilization of Ethiopian wolves (Canis simensis). J Wildl Dis, 1996. 32(1): p. 147-51.


Sillero-Zubiri, C., A.A. King, and D.W. Macdonald, Rabies and mortality in Ethiopian wolves (Canis simensis). J Wildl Dis, 1996. 32(1): p. 80-6.


Sillero-Zubiri, C., D. Gottelli, and D.W. Macdonald, Male philopatry, extra-pack copulations and inbreeding avoidance in Ethiopian wolves ( Canis simensis ). Behavioral Ecology and Sociobiology, 1996. 38(5): p. 331-340. Link


Sillero-Zubiri, C. and D. Gottelli, Diet and Feeding Behavior of Ethiopian Wolves (Canis simensis). Journal of Mammalogy, 1995. 76(2): p. 531-541.


Sillero-Zubiri, C. and D. Gottelli, Spatial organization in the Ethiopian wolf Canis simensis: Large packs and small stable home ranges. Journal of Zoology (London), 1995. 237(1): p. 65-81.


Sillero-Zubiri, C., F.H. Tattersall, and D.W. Macdonald, Bale Mountains rodent communities and their relevance to the Ethiopian wolf (Canis simensis). African Journal of Ecology, 1995. 33(4): p. 301-320.


Sillero-Zubiri, C., F.H. Tattersall, and D.W. Macdonald, Habitat selection and daily activity of giant molerats Tachyoryctes macrocephalus: Significance to the Ethiopian wolf Canis simensis in the Afroalpine ecosystem. Biological Conservation, 1995. 72(1): p. 77-84. Link


Gottelli, D., et al., Molecular genetics of the most endangered canid: The Ethiopian wolf Canis simensis. Molecular Ecology, 1994. 3(4): p. 301-312.


Gottelli, D. and C. Sillero-Zubiri, The Ethiopian wolf - an endangered endemic canid. Oryx, 1992. 26: p. 205-214. 


Mebatsion, T., et al., Detection of rabies antibody by ELISA and RFFIT in unvaccinated dogs and in the endangered Simien jackal (Canis simensis) of Ethiopia. Zentralbl Veterinarmed B, 1992. 39(3): p. 233-5.